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Noun quill has 4 senses
quill, quill pen - pen made from a bird's feather--1 is a kind of pen
 
quill - a stiff hollow protective spine on a porcupine or hedgehog--2 is a kind of spine
 --2 is a part of
 flight feather, pinion, quill, quill feather - any of the larger wing or tail feathers of a bird--3 is a kind of feather, plume, plumage
 --3 is a part of wing
 --3 has particulars: primary, primary feather, primary quill; tail feather
 
quill, calamus, shaft - the hollow shaft of a feather--4 is a kind of rib
 --4 is a part of feather, plume, plumage
 
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